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(Ooki et al, 1990, 1993)\u3002\u4eca\u56de\u3001\u5bfe\u8c61\u3092\u5927\u5e45\u306b\u5897\u52a0\u3057\u3001\u307e\u305f\u65b0\u305f\u306a\u8cea\u554f\u9805\u76ee\u3092\u8ffd\u52a0\u3059\u308b\u3053\u3068\u3067\u5375\u6027\u8a3a\u65ad\u7528\u8cea\u554f\u7d19\u7968\u306e\u518d\u691c\u8a0e\u3092\u884c\u3063\u305f\u3002\uff0f\u5bfe\u8c61\u3000\u5bfe\u8c61\u306f\u30011985\u5e74\u5ea6\u304b\u30892003\u5e74\u5ea6\u307e\u3067\u306b\u6771\u4eac\u5927\u5b66\u6559\u80b2\u5b66\u90e8\u9644\u5c5e\u4e2d\u7b49\u6559\u80b2\u5b66\u6821\u306b\u5165\u5b66\u5fd7\u9858\u306a\u3044\u3057\u5165\u5b66\u3057\u305f\u540c\u6027\u53cc\u751f\u5150224\u7d44\u3067\u3042\u308b\u3002\u5150\u306e\u5e74\u9f62\u306f11\u6b73\u306a\u3044\u305712\u6b73\u3067\u3042\u308b\u3002\u540c\u6821\u5165\u5b66\u5fd7\u9858\u6642\u306b\u63d0\u51fa\u3055\u308c\u308b\u53cc\u751f\u5150\u8abf\u67fb\u7968\u306e\u4e2d\u306b\u4eca\u56de\u306e\u5206\u6790\u306b\u5229\u7528\u3057\u305f\u5375\u6027\u8a3a\u65ad\u7528\u8cea\u554f\u7d19\u7968\u304c\u542b\u307e\u308c\u3066\u3044\u308b\u3002\u4fdd\u8b77\u8005\uff08\u4e3b\u3068\u3057\u3066\u6bcd\u89aa\uff09\u306f\u3053\u308c\u3089\u306e\u8cc7\u6599\u3092\u3082\u3068\u306b\u533b\u5b66\u9762\u63a5\u3092\u53d7\u3051\u3001\u5408\u308f\u305b\u3066\u305d\u306e\u8a18\u8f09\u306e\u78ba\u8a8d\u3082\u884c\u308f\u308c\u308b\u3002\u7b46\u8005\u306f\u3001\u4e0a\u8a18\u306e\u5e74\u5ea6\u5168\u3066\u306b\u304a\u3044\u3066\u9762\u63a5\u8005\u306e\u4e00\u4eba\u3068\u3057\u3066\u7acb\u3061\u4f1a\u3063\u3066\u3044\u308b\u3002\u5fd7\u9858\u8005\u306e\u3046\u3061\u6bcd\u5b50\u624b\u5e33\u7b49\u306b\u3088\u308a\u8840\u6db2\u578b\u4e0d\u4e00\u81f4\u304c\u78ba\u8a8d\u3055\u308c\u305f\u3082\u306e\u306f\u4e8c\u5375\u6027\u3068\u8a3a\u65ad\u3055\u308c\u308b\u3002\u5165\u5b66\u8005\u306b\u5bfe\u3057\u3066\u306fDNA\u691c\u67fb\u3092\u542b\u3081\u305f\u591a\u6570\u306e\u907a\u4f1d\u30de\u30fc\u30ab\u30fc\u306b\u3088\u308a\u5375\u6027\u3092\u6c7a\u5b9a\u3057\u3066\u3044\u308b\u3002\u305d\u306e\u7d50\u679c\u3001\u5206\u6790\u5bfe\u8c61\u306e\u5375\u6027\u306e\u5185\u8a33\u306f\u4e00\u5375\u6027159\u7d44\u3001\u4e8c\u5375\u602765\u7d44\u3068\u306a\u3063\u305f\u3002\u3053\u306e\u5375\u6027\u5225\u983b\u5ea6\u306f\u4eba\u53e3\u52d5\u614b\u7d71\u8a08\u306b\u57fa\u3065\u304d\u63a8\u5b9a\u3055\u308c\u305f\u3001\u308f\u304c\u56fd\u306e\u5375\u6027\u5225\u983b\u5ea6\u3068\u5927\u5dee\u306f\u306a\u3044\u3002\u4ee5\u4e0a\u306e\u691c\u67fb\u306f\u5165\u8a66\u306e\u4e00\u8cab\u3068\u3057\u3066\u5b9f\u65bd\u3055\u308c\u3001\u66f8\u9762\u306b\u3088\u308b\u540c\u610f\u306e\u3082\u3068\u306b\u5b9f\u65bd\u3055\u308c\u3066\u3044\u308b\u3002\uff0f\u65b9\u6cd5\uff0f\u5375\u6027\u8a3a\u65ad\u7528\u8cea\u554f\u7d19\u7968\u3000\u6bcd\u89aa\u7528\u8cea\u554f\u7d19\u7968\u306f\u53cc\u751f\u5150\u304c\u304a\u3088\u305d1\u6b73\u306e\u3053\u308d\u306e\u5168\u4f53\u7684\u306a\u985e\u4f3c\u5ea6\u306b\u95a2\u3059\u308b3\u3064\u306e\u8cea\u554f\u9805\u76ee (Ooki et al, 1993) \u3068\u3001\u8eab\u4f53\u7684\u985e\u4f3c\u306b\u95a2\u3059\u308b16\u306e\u8cea\u554f\u9805\u76ee\u304b\u3089\u306a\u308b\u30023\u3064\u306e\u8cea\u554f\u9805\u76ee\u306f\u3001\u300c\u4e8c\u4eba\u306f\u74dc\u4e8c\u3064\u306e\u3088\u3046\u306b\u4f3c\u3066\u3044\u305f\u304b\u300d\u3001\u300c\u3075\u305f\u308a\u306f\u3057\u3070\u3057\u3070\u9593\u9055\u3048\u3089\u308c\u305f\u304b\u300d\u3001\u300c\u8ab0\u306b\u9593\u9055\u3048\u3089\u308c\u305f\u304b\u300d\u3067\u3042\u308b\u3002\u8eab\u4f53\u7684\u306a\u985e\u4f3c\u306b\u95a2\u3059\u308b\u8cea\u554f\u9805\u76ee\u306f\u3001\u300c\u9854\u306e\u304b\u305f\u3061\u300d\u3001\u300c\u8033\u306e\u304b\u305f\u3061\u300d\u3001\u300c\u7709\u306e\u304b\u305f\u3061\u300d\u3001\u300c\u6307\u306e\u304b\u305f\u3061\u300d\u3001\u300c\u5bdd\u304c\u304a\u300d\u3001\u300c\u5bdd\u305e\u3046\u300d\u3001\u300c\u58f0\u300d\u306a\u3069\u3067\u3042\u308b\u3002\u53cc\u751f\u5150\u7528\u306e\u8cea\u554f\u7d19\u7968\u306f\u73fe\u5728\u5e74\u9f62\u306b\u304a\u3051\u308b\u5168\u4f53\u7684\u306a\u985e\u4f3c\u5ea6\u306b\u95a2\u3059\u308b3\u3064\u306e\u8cea\u554f\u9805\u76ee (Ooki et al, 1990) \u3060\u3051\u304b\u3089\u306a\u308a\u3001\u53cc\u751f\u5150\u4e8c\u4eba\u306b\u5225\u3005\u306b\u56de\u7b54\u3057\u3066\u3082\u3089\u3046\u3002\u8cea\u554f\u9805\u76ee\u306f\u6bcd\u89aa\u7528\u306e\u8cea\u554f\u3068\u540c\u69d8\u306b\u3001\u300c\u4e8c\u4eba\u306f\u74dc\u4e8c\u3064\u306e\u3088\u3046\u306b\u4f3c\u3066\u3044\u305f\u304b\u300d\u3001\u300c\u3075\u305f\u308a\u306f\u3057\u3070\u3057\u3070\u9593\u9055\u3048\u3089\u308c\u305f\u304b\u300d\u3001\u300c\u8ab0\u306b\u9593\u9055\u3048\u3089\u308c\u305f\u304b\u300d\u306e3\u9805\u76ee\u3067\u3042\u308b\u3002\u305d\u308c\u305e\u308c\u306e\u8cea\u554f\u306b\u5bfe\u3059\u308b\u56de\u7b54\u306b\u306f\u985e\u4f3c\u306e\u7a0b\u5ea6\u306b\u3088\u3063\u30661\u70b9\u304b\u30893\u70b9\uff08\u300c\u8ab0\u306b\u9593\u9055\u3048\u3089\u308c\u305f\u304b\u300d\u306e\u307f1\u70b9\u304b\u30894\u70b9\uff09\u304c\u4e0e\u3048\u3089\u308c\u308b\uff08\u70b9\u6570\u304c\u4f4e\u3044\u7a0b\u985e\u4f3c\u5ea6\u304c\u9ad8\u3044\uff09\u3002\u3053\u306e\u56de\u7b54\u7d50\u679c\u3092\u57fa\u306b\u5375\u6027\u3092\u5224\u5b9a\u3059\u308b\u3002\uff0f\u7d71\u8a08\u89e3\u6790\u3000\u907a\u4f1d\u30de\u30fc\u30ab\u30fc\u306b\u3088\u308b\u5375\u6027\u3068\u8cea\u554f\u7d19\u306b\u3088\u308b\u5375\u6027\u306e\u8a3a\u65ad\u7d50\u679c\u3092\u6bd4\u8f03\u3057\u8cea\u554f\u7d19\u7968\u306e\u6709\u52b9\u6027\u3092\u691c\u8a0e\u3057\u305f\u3002\u8a3a\u65ad\u306e\u6b63\u78ba\u5ea6\u5224\u5b9a\u306b\u5f53\u305f\u3063\u3066\u306f\u907a\u4f1d\u30de\u30fc\u30ab\u30fc\u306b\u3088\u308a\u6c7a\u5b9a\u3057\u305f\u5375\u6027\u3092\u5f93\u5c5e\u5909\u6570\u3001\u8cea\u554f\u7d19\u7968\u306e\u5404\u9805\u76ee\u3092\u8aac\u660e\u5909\u6570\u3068\u3059\u308b\u3001\u5909\u6570\u9078\u629e\u6cd5\u591a\u91cd\u30ed\u30b8\u30b9\u30c6\u30a3\u30c3\u30af\u5206\u6790\u3092\u5b9f\u65bd\u3057\u305f\u3002\u305d\u306e\u969b\u3001\u6bcd\u89aa\u7528\u306e\u8cea\u554f\u7d19\u3067\u306f\u3001\u5168\u4f53\u7684\u306a\u985e\u4f3c\u306b\u95a2\u3059\u308b\u8cea\u554f\u9805\u76ee\u306e\u307f\u3001\u8eab\u4f53\u7684\u985e\u4f3c\u306b\u95a2\u3059\u308b\u8cea\u554f\u9805\u76ee\u306e\u307f\u3001\u4e21\u8005\u3092\u3059\u3079\u3066\u5408\u308f\u305b\u305f\u8cea\u554f\u9805\u76ee\u306b\u3088\u308b\u5206\u6790\u3092\u5b9f\u65bd\u3057\u305f\u3002\u53cc\u751f\u5150\u306b\u95a2\u3057\u3066\u306f\u3001\u4e8c\u4eba\u306e\u56de\u7b54\u7d50\u679c\u3092\u5229\u7528\u3057\u305f\u5834\u5408\u3068\u3069\u3061\u3089\u304b\u4e00\u65b9\u306e\u56de\u7b54\u306e\u307f\u3092\u5229\u7528\u3057\u305f\u5834\u5408\u306e\u5206\u6790\u3092\u884c\u3063\u305f\u3002\u307e\u305f\u3001\u3053\u306e\u8cea\u554f\u7d19\u7968\u306e\u4e00\u822c\u7684\u306a\u666e\u53ca\u3092\u8003\u616e\u3057\u3066\u3001\u5f93\u6765\u3069\u304a\u308a\u306b\u8cea\u554f\u7d19\u306e\u56de\u7b54\u7d50\u679c\u306e\u5358\u7d14\u548c\u306b\u3088\u308b\u5375\u6027\u306e\u63a8\u5b9a\u3092\u5408\u308f\u305b\u3066\u884c\u3063\u305f\u3002\u3053\u306e\u5834\u5408\u3001\u8a3a\u65ad\u306b\u6709\u52b9\u3068\u601d\u308f\u308c\u308b\u6570\u9805\u76ee\u306e\u56de\u7b54\u306e\u5358\u7d14\u306a\u5408\u8a08\u5f97\u70b9\u306e\u5206\u5e03\u3092\u3082\u3068\u306b\u3001\u9069\u5f53\u306a\u30ab\u30c3\u30c8\u30aa\u30d5\u30dd\u30a4\u30f3\u30c8\u3092\u8a2d\u5b9a\u3057\u5375\u6027\u3092\u5224\u5b9a\u3059\u308b\u3002\u4f8b\u3048\u3070\u3001\u5f93\u6765\u306e\u6bcd\u89aa\u7528\u8cea\u554f\u7d19\u7968 (Ooki et al, 1993) \u3067\u306f\u30013\u9805\u76ee\u3068\u3082\u306b1\u70b9\u306e\u5834\u5408\u306e3\u70b9\u304b\u3089\u3001\u305d\u308c\u305e\u308c\u306e\u8cea\u554f\u306b3\u30013\u30014\u3068\u56de\u7b54\u3057\u305f\u5834\u5408\u306e10\u70b9\u307e\u3067\u306b\u5206\u5e03\u3059\u308b\u3002\u3053\u306e\u5834\u5408\u30016\u70b9\u4ee5\u4e0b\u3092\u4e00\u5375\u6027\u3068\u30017\u70b9\u4ee5\u4e0a\u3092\u4e8c\u5375\u6027\u3068\u3059\u308b\u3068\u6b63\u78ba\u5ea6\u304c\u304a\u3088\u305d90\uff05\u7a0b\u5ea6\u3068\u306a\u308b\u3002\uff0f\u7d50\u679c\uff0f\u591a\u91cd\u30ed\u30b8\u30b9\u30c6\u30a3\u30c3\u30af\u5206\u6790\u306e\u7d50\u679c\u3000\u6bcd\u89aa\u7528\u8cea\u554f\u7d19\u306b\u304a\u3051\u308b\u8eab\u4f53\u7684\u985e\u4f3c\u5ea6\u306b\u95a2\u3059\u308b16\u9805\u76ee\u3092\u7528\u3044\u305f\u5834\u5408\u306e\u8a3a\u65ad\u306e\u6b63\u78ba\u5ea6\u306f91.5\uff05\u3001\u5168\u4f53\u7684\u985e\u4f3c\u5ea6\u306b\u95a2\u3059\u308b3\u9805\u76ee\u3092\u7528\u3044\u305f\u5834\u5408\u306e\u8a3a\u65ad\u306e\u6b63\u78ba\u5ea6\u306f91.5\uff05\u3067\u3042\u3063\u305f\u3002\u516819\u9805\u76ee\u3092\u7528\u3044\u305f\u5834\u5408\u306e\u8a3a\u65ad\u306e\u6b63\u78ba\u5ea6\u306f95.1\uff05\u3067\u3042\u308a\u3001\u4e00\u5375\u6027\u306e96.2\uff05\u3001\u4e8c\u5375\u6027\u306e92.3\uff05\u304c\u6b63\u3057\u304f\u5224\u5b9a\u3055\u308c\u305f\u3002\u3053\u306e\u5834\u5408\u300c\u4e8c\u4eba\u304c\u9593\u9055\u3048\u3089\u308c\u305f\u983b\u5ea6\u300d\u3001\u300c\u6307\u306e\u304b\u305f\u3061\u300d\u3001\u300c\u7709\u306e\u304b\u305f\u3061\u300d\u304c\u6709\u610f\u306a\u9805\u76ee\u3068\u3057\u3066\u63a1\u629e\u3055\u308c\u305f\u3002\u53cc\u751f\u5150\u672c\u4eba\u7528\u8cea\u554f\u7d19\u7968\u306b\u3088\u308b\u6b63\u78ba\u5ea6\u306f93.3\uff05\u3067\u3042\u3063\u305f\u3002\u53cc\u751f\u5150\u4e00\u4eba\u306e\u56de\u7b54\u3092\u5229\u7528\u3057\u305f\u5834\u5408\u306b\u306f\u6b63\u78ba\u5ea6\u306f92.0\uff05\u3067\u3042\u3063\u305f\u3002\u6bcd\u89aa\u7528\u8cea\u554f\u7d19\u7968\u3068\u53cc\u751f\u5150\u672c\u4eba\u7528\u8cea\u554f\u7d19\u7968\u306e\u56de\u7b54\u3092\u540c\u6642\u306b\u5206\u6790\u3057\u3066\u3082\u6b63\u78ba\u5ea6\u306e\u5927\u5e45\u306a\u6539\u5584\u306f\u898b\u3089\u308c\u306a\u304b\u3063\u305f\u3002\uff0f\u8cea\u554f\u7d19\u306e\u56de\u7b54\u306e\u5358\u7d14\u548c\u306b\u3088\u308b\u5206\u6790\u306e\u7d50\u679c\u3000\u5375\u6027\u306b\u3088\u308b\u5358\u7d14\u548c\u306e\u5206\u5e03\u306e\u5dee\u306f\u660e\u77ad\u3067\u3042\u3063\u305f\u3002\u4e00\u5375\u6027\u3067\u306f\u5408\u8a08\u5f97\u70b9\u306f\u5c0f\u3055\u3044\u5024\uff08\u985e\u4f3c\u5ea6\u304c\u9ad8\u3044\uff09\u306b\u5206\u5e03\u3057\u3001\u4e8c\u5375\u6027\u3067\u306f\u5408\u8a08\u5f97\u70b9\u306f\u5927\u304d\u3044\u5024\uff08\u985e\u4f3c\u5ea6\u304c\u4f4e\u3044\uff09\u306b\u5206\u5e03\u3057\u305f\u3002\u305f\u3060\u3057\u3001\u5206\u5e03\u306b\u306f\u304b\u306a\u308a\u306e\u30aa\u30fc\u30d0\u30fc\u30e9\u30c3\u30d7\u304c\u8a8d\u3081\u3089\u308c\u305f\u3002\u6bcd\u89aa\u7528\u8cea\u554f\u7d19\u7968\u306b\u5bfe\u3057\u3066 Ooki et al (1993) \u306e\u5224\u5b9a\u57fa\u6e96\uff08\u5168\u4f53\u7684\u306a\u985e\u4f3c\u306b\u95a2\u3059\u308b3\u9805\u76ee\u306e\u56de\u7b54\u306e\u5358\u7d14\u548c\uff09\u3092\u7528\u3044\u308b\u3068\u6b63\u78ba\u5ea6\u306f89.7\uff05\u3067\u3042\u308a\u3001\u53cc\u751f\u5150\u672c\u4eba\u7528\u306b\u5bfe\u3057\u3066 Ooki et al (1990) 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(2.00-3.66) 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