{"_buckets": {"deposit": "beafbdf5-6070-4d9a-b7b4-7498db230ee6"}, "_deposit": {"id": "4310", "owners": [], "pid": {"revision_id": 0, "type": "depid", "value": "4310"}, "status": "published"}, "_oai": {"id": "oai:repository.dl.itc.u-tokyo.ac.jp:00004310"}, "item_7_alternative_title_1": {"attribute_name": "\u305d\u306e\u4ed6\u306e\u30bf\u30a4\u30c8\u30eb", "attribute_value_mlt": [{"subitem_alternative_title": "Evolution of genes duplicated through fish-specific genome doubling"}]}, "item_7_biblio_info_7": {"attribute_name": "\u66f8\u8a8c\u60c5\u5831", "attribute_value_mlt": [{"bibliographicIssueDates": {"bibliographicIssueDate": "2008-03-24", "bibliographicIssueDateType": "Issued"}, "bibliographic_titles": [{}]}]}, "item_7_date_granted_25": {"attribute_name": "\u5b66\u4f4d\u6388\u4e0e\u5e74\u6708\u65e5", "attribute_value_mlt": [{"subitem_dategranted": "2008-03-24"}]}, "item_7_degree_grantor_23": {"attribute_name": "\u5b66\u4f4d\u6388\u4e0e\u6a5f\u95a2", "attribute_value_mlt": 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genome doubling, FSGD)\u3092\u7d4c\u9a13\u3057\u3066\u3044\u308b\u9b5a\u985e\u306b\u7740\u76ee\u3057\u305f.FSGD\u306b\u3088\u3063\u3066\u91cd\u8907\u3057\u305f\u907a\u4f1d\u5b50\u306f,\u73fe\u5728\u306e\u9b5a\u985e\u30b2\u30ce\u30e0\u4e2d\u306b\u591a\u304f\u5b58\u7d9a\u3057\u3066\u3044\u308b\u3068\u671f\u5f85\u3055\u308c\u308b\u304c,\u8a73\u7d30\u306f\u3088\u304f\u308f\u304b\u3063\u3066\u3044\u306a\u3044.\u305d\u3053\u3067\u5168\u30b2\u30ce\u30e0\u914d\u5217\u304c\u6c7a\u5b9a\u3055\u308c\u305f\u9b5a\u985e4\u7a2e(\u30bc\u30d6\u30e9\u30d5\u30a3\u30c3\u30b7\u30e5,\u30e1\u30c0\u30ab,\u30a4\u30c8\u30e8,\u30df\u30c9\u30ea\u30d5\u30b0)\u306b\u7740\u76ee\u3057,\u305d\u308c\u3089\u306e\u30b2\u30ce\u30e0\u306e\u907a\u4f1d\u5b50\u30ec\u30d1\u30fc\u30c8\u30ea\u30fc\u3092,FSGD\u3092\u7d4c\u3066\u3044\u306a\u3044\u56db\u8db3\u985e\u30b2\u30ce\u30e0\u306e\u907a\u4f1d\u5b50\u30ec\u30d1\u30fc\u30c8\u30ea\u30fc\u3068\u6bd4\u8f03\u691c\u8a0e\u3059\u308b\u3053\u3068\u3067,FSGD\u306b\u3088\u3063\u3066\u91cd\u8907\u3057\u305f\u907a\u4f1d\u5b50\u7fa4\u304c,\u305d\u306e\u5f8c\u306b\u3069\u306e\u3088\u3046\u306a\u6b20\u5931\u307e\u305f\u306f\u5b58\u7d9a\u904e\u7a0b\u3092\u7d4c\u3066,\u73fe\u5728\u306e\u9b5a\u985e\u30b2\u30ce\u30e0\u306b\u3069\u306e\u7a0b\u5ea6\u7dad\u6301\u3055\u308c\u3066\u3044\u308b\u306e\u304b,\u307e\u305f,\u7dad\u6301\u3055\u308c\u3066\u304d\u305f\u91cd\u8907\u907a\u4f1d\u5b50\u304c\u6a5f\u80fd\u5206\u5316\u3092\u7d4c\u3066\u304d\u305f\u306e\u304b\u3092\u691c\u8a0e\u3057\u305f.\u6b21\u306b,FSGD\u306b\u7531\u6765\u3059\u308b\u3068\u8003\u3048\u3089\u308c\u308b\u91cd\u8907\u907a\u4f1d\u5b50Pgi\u306b\u7740\u76ee\u3057,\u305d\u308c\u3089\u304c\u30b3\u30fc\u30c9\u3059\u308b\u30bf\u30f3\u30d1\u30af\u8cea(phosphoglucose isomerase, EC 5.3.1.9)\u306e\u65b0\u3057\u3044\u7279\u6027,\u3068\u304f\u306b\u96fb\u8377\u304c\u9032\u5316\u3057\u3066\u304d\u305f\u904e\u7a0b\u3092,\u9b5a\u985e\u306e\u7cfb\u7d71\u67a0\u306b\u7acb\u811a\u3057\u3066\u8a73\u7d30\u306b\u89e3\u6790\u3057\u305f.\u3055\u3089\u306b,Pgi\u306e\u89e3\u6790\u304b\u3089\u660e\u3089\u304b\u306b\u306a\u3063\u305f\u5206\u5b50\u9032\u5316\u69d8\u5f0f\u306e\u666e\u904d\u6027\u3092\u691c\u8a0e\u3059\u308b\u305f\u3081\u306b,\u810a\u690e\u52d5\u7269\u306b\u7279\u7570\u7684\u306a\u91cd\u8907\u907a\u4f1d\u5b50Ald\u3092\u89e3\u6790\u3057\u305f.\u3053\u308c\u3089\u306e\u7d50\u679c\u3092\u6bd4\u8f03\u691c\u8a0e\u3057,\u65b0\u3057\u3044\u30bf\u30f3\u30d1\u30af\u8cea\u7279\u6027\u304c\u9032\u5316\u3059\u308b\u969b\u306e\u5206\u5b50\u9032\u5316\u69d8\u5f0f\u3068\u9078\u629e\u5727\u306e\u5f37\u3055\u3068\u306e\u95a2\u9023\u3092\u8b70\u8ad6\u3057\u305f.1.\u9b5a\u985e\u7279\u7570\u7684\u30b2\u30ce\u30e0\u500d\u5316\u306b\u7531\u6765\u3059\u308b\u91cd\u8907\u907a\u4f1d\u5b50\u306e\u5b58\u7d9a\u3068\u9032\u5316FSGD\u306b\u3088\u3063\u3066\u91cd\u8907\u3057,\u305d\u306e\u5f8c\u73fe\u5728\u307e\u3067\u9b5a\u985e\u30b2\u30ce\u30e0\u4e2d\u306b\u5b58\u7d9a\u3057\u3066\u304d\u305f\u907a\u4f1d\u5b50\u306e\u5272\u5408\u3092\u63a8\u5b9a\u3059\u308b\u305f\u3081\u306b,\u8a18\u61b6\u3068\u5b66\u7fd2,\u74b0\u5883\u60c5\u5831\u51e6\u7406\u304a\u3088\u3073\u57fa\u790e\u4ee3\u8b1d\u306b\u95a2\u308f\u308b\u907a\u4f1d\u5b50\u7fa4\u306b\u7740\u76ee\u3057,\u305d\u308c\u3089\u304b\u3089,\u30b7\u30ca\u30d7\u30b9\u4f1d\u9054\u306e\u9577\u671f\u5897\u5f37,\u5473\u899a\u4f1d\u9054,\u55c5\u899a\u4f1d\u9054,\u30af\u30a8\u30f3\u9178\u56de\u8def\u306b\u95a2\u4e0e\u3059\u308b130\u500b\u306e\u30d2\u30c8\u907a\u4f1d\u5b50\u3092\u89e3\u6790\u306e\u7acb\u811a\u70b9\u3068\u3057\u305f.\u3053\u308c\u3089\u306b\u76f8\u540c\u306a\u907a\u4f1d\u5b50\u3092,\u30bc\u30d6\u30e9\u30d5\u30a3\u30c3\u30b7\u30e5,\u30e1\u30c0\u30ab,\u30a4\u30c8\u30e8,\u30df\u30c9\u30ea\u30d5\u30b0,\u30cb\u30ef\u30c8\u30ea,\u30cd\u30c3\u30bf\u30a4\u30c4\u30e1\u30ac\u30a8\u30eb,\u30ab\u30bf\u30e6\u30a6\u30ec\u30a4\u30dc\u30e4,\u30ad\u30a4\u30ed\u30b7\u30e7\u30a6\u30b8\u30e7\u30a6\u30d0\u30a8\u306e\u5168\u30b2\u30ce\u30e0\u914d\u5217\u304b\u3089\u63a2\u7d22\u3057\u305f.\u5f97\u3089\u308c\u305f\u30c7\u30fc\u30bf\u3092\u3082\u3068\u306b,\u6700\u5c24\u6cd5\u306b\u3088\u308b\u53b3\u5bc6\u306a\u5206\u5b50\u7cfb\u7d71\u89e3\u6790\u3092\u500b\u3005\u306e\u907a\u4f1d\u5b50\u30d5\u30a1\u30df\u30ea\u30fc\u306b\u3064\u3044\u3066\u884c\u3063\u305f\u7d50\u679c,130\u500b\u306e\u30d2\u30c8\u907a\u4f1d\u5b50\u306e\u3046\u306114\u500b\u306f\u30d2\u30c8\u3082\u3057\u304f\u306f\u56db\u8db3\u985e\u306b\u7279\u7570\u7684\u3067\u3042\u3063\u305f\u304c,\u6b8b\u308a\u306e116\u500b\u3067\u9b5a\u985e\u306e\u76f8\u540c\u907a\u4f1d\u5b50\u304c\u540c\u5b9a\u3055\u308c\u305f.\u540c\u5b9a\u3055\u308c\u305f\u76f8\u540c\u907a\u4f1d\u5b50\u7fa4\u306e\u7cfb\u7d71\u95a2\u4fc2\u304b\u3089,FSGD\u306b\u3088\u308b\u91cd\u8907\u30a4\u30d9\u30f3\u30c8\u304c45\u500b\u691c\u51fa\u3055\u308c\u305f.\u9b5a\u985e\u306e\u63a8\u5b9a\u5206\u5c90\u5e74\u4ee3\u3068\u4f75\u305b\u3066\u691c\u8a0e\u3059\u308b\u3068,FSGD\u304c\u8d77\u304d\u3066\u304b\u3089\u4e0a\u8a18\u306e\u9b5a\u985e\u306e\u5171\u901a\u7956\u5148\u304c\u767b\u5834\u3059\u308b\u307e\u3067\u306e\u7d043\u5343\u4e07\u5e74\u306e\u9593\u306b,\u91cd\u8907\u907a\u4f1d\u5b50\u306e61.2% (71/116) \u304c\u5931\u308f\u308c\u3066\u304d\u305f\u4e00\u65b9\u3067,\u3053\u306e\u5171\u901a\u7956\u5148\u30b2\u30ce\u30e0\u306b\u6b8b\u3063\u3066\u3044\u305f\u91cd\u8907\u907a\u4f1d\u5b50\u306e\u5e73\u574761.1% (27.5/45) \u304c,\u305d\u306e\u5f8c\u7d043\u5104\u5e74\u9593\u5b58\u7d9a\u3057\u3066\u73fe\u5728\u306e\u9b5a\u985e\u30b2\u30ce\u30e0\u306b\u542b\u307e\u308c\u3066\u3044\u308b\u3053\u3068\u304c\u308f\u304b\u3063\u305f.\u3053\u306e\u3053\u3068\u306f,FSGD\u306e\u76f4\u5f8c\u306f\u91cd\u8907\u907a\u4f1d\u5b50\u304c\u6025\u901f\u306b\u5931\u308f\u308c\u308b\u4e00\u65b9\u3067,\u6b8b\u3063\u305f\u91cd\u8907\u907a\u4f1d\u5b50\u306f\u6bd4\u8f03\u7684\u9577\u671f\u9593,\u30b2\u30ce\u30e0\u306b\u7dad\u6301\u3055\u308c\u3066\u304d\u305f\u3053\u3068\u3092\u793a\u5506\u3059\u308b.FSGD\u3067\u91cd\u8907\u3057\u305f\u5f8c\u306b\u9577\u671f\u9593\u7dad\u6301\u3055\u308c\u3066\u304d\u305f\u907a\u4f1d\u5b50\u7fa4\u304c\u3069\u306e\u3088\u3046\u306a\u7279\u6027\u3092\u3082\u3064\u3082\u306e\u3067\u3042\u308b\u306e\u304b\u3092,\u30b3\u30fc\u30c9\u3059\u308b\u30bf\u30f3\u30d1\u30af\u8cea\u306b\u7740\u76ee\u3057\u3066\u89e3\u6790\u3057\u305f\u3068\u3053\u308d,\u30bf\u30f3\u30d1\u30af\u8cea\u306e\u30b5\u30a4\u30ba,\u304a\u3088\u3073\u76f8\u4e92\u4f5c\u7528\u30d1\u30fc\u30c8\u30ca\u30fc\u6570\u304c,FSGD\u306e\u76f4\u5f8c\u306b\u5931\u308f\u308c\u305f\u91cd\u8907\u907a\u4f1d\u5b50\u7fa4\u3068\u6bd4\u3079\u3066\u6709\u610f\u306b\u9ad8\u3044\u5024\u3092\u793a\u3057\u305f(\u524d\u8005: \u5927\u30fb\u4e2d\u30fb\u5c0f3\u7fa4\u9593\u306e x2 test, P=0.0052; \u5f8c\u8005: Welch\u0026apos;s t test, P=0.0288).\u30bf\u30f3\u30d1\u30af\u8cea\u306e\u30b5\u30a4\u30ba(\u5927\u304d\u306a\u30bf\u30f3\u30d1\u30af\u8cea\u307b\u3069\u591a\u304f\u306e\u30c9\u30e1\u30a4\u30f3\u3084\u30e2\u30c1\u30fc\u30d5\u3092\u3082\u3064\u3068\u671f\u5f85\u3055\u308c\u308b),\u304a\u3088\u3073\u76f8\u4e92\u4f5c\u7528\u30d1\u30fc\u30c8\u30ca\u30fc\u6570\u306f,\u907a\u4f1d\u5b50\u306e\u591a\u6a5f\u80fd\u6027\u3092\u53cd\u6620\u3057\u3066\u3044\u308b\u3068\u8003\u3048\u3089\u308c\u308b.\u3053\u306e\u3053\u3068\u304b\u3089,\u3042\u3089\u304b\u3058\u3081\u591a\u304f\u306e\u6a5f\u80fd\u3092\u3082\u3063\u3066\u3044\u305f\u907a\u4f1d\u5b50\u304c,FSGD\u306b\u3088\u3063\u3066\u91cd\u8907\u3057\u305f\u5f8c\u306b\u3088\u308a\u9ad8\u3044\u78ba\u7387\u3067\u6a5f\u80fd\u5206\u5316\u3092\u7d4c\u305f\u7d50\u679c,\u9577\u671f\u9593\u7dad\u6301\u3055\u308c\u3066\u304d\u305f\u3082\u306e\u3068\u8003\u3048\u3089\u308c\u305f.\u3053\u306e\u7d50\u679c\u306f,\u91cd\u8907\u907a\u4f1d\u5b50\u304c\u9032\u5316\u7684\u306b\u5b58\u7d9a\u3059\u308b\u4e00\u6b21\u904e\u7a0b\u3068\u3057\u3066,\u6a5f\u80fd\u5206\u5316\u304c\u91cd\u8981\u306a\u5f79\u5272\u3092\u679c\u305f\u3059\u3053\u3068\u3092\u793a\u3059\u65b0\u305f\u306a\u8a3c\u62e0\u3092\u63d0\u4f9b\u3059\u308b.FSGD\u3067\u91cd\u8907\u3057,\u6a5f\u80fd\u5206\u5316\u3092\u7d4c\u3066\u7dad\u6301\u3055\u308c\u3066\u304d\u305f\u907a\u4f1d\u5b50\u304c,\u305d\u306e\u5f8c,\u7cfb\u7d71\u7279\u7570\u7684\u306b\u65b0\u6a5f\u80fd\u3092\u7372\u5f97\u3059\u308b\u3053\u3068\u3067\u9b5a\u985e\u9593\u306e\u907a\u4f1d\u7684\u5dee\u7570\u3092\u3082\u305f\u3089\u3057,\u9b5a\u985e\u304c\u793a\u3059\u591a\u69d8\u6027\u306b\u3055\u307e\u3056\u307e\u306a\u5f62\u3067\u95a2\u4e0e\u3057\u3066\u304d\u305f\u3053\u3068\u304c\u63a8\u5bdf\u3055\u308c\u308b.2.\u9b5a\u985e\u7279\u7570\u7684\u306a\u91cd\u8907\u907a\u4f1d\u5b50 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\u524d\u8005: 6.21-6.36; \u5f8c\u8005: 6.75-7.36; U test, 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